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You know how sometimes in the Vault if Glass you'll see what looks like little bits if dust? Even when yiu fight the Nexus, you see them. If you get close enough you'll see that they are actually Ghosts without a shell. Phantom Ghosts, whose Guardians were lost in the darkest corners of time. They were born to protect their homes and failed.
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Durn. I'll think a little more on this then.