There are two scenarios: worms = ahamkara or worms parasite have infected other species including ahamkara
The "oh bearer of mine" is a dead give away. The description of Yul is very dragon like.
Yet in the "id shut them all in cells.." card about dragons, he says our gods should be ours alone. Which could easily mean the dragon (ahamkara) contains the same worm parasite (God) as the hive.
To say worm = ahamkara would imply that in the past guardians killed some of the hive gods, because we defeated the ahamkara. But yet Toland is trying to figure out the names of the hive gods after that happened. If we fought Yul or eir or akka or some other unnamed ahamkara, we wouldn't be so unsure about the hive or its origins, at least I think so
I think all the book of sorrows says is that the worms, whose true nature and species is not known, commune directly with the darkness, and their parasite have infected at least the hive and ahamkara.
I think there is interpretation for both. But you guys know a lot more than I on this
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