I just replayed the 1st mission again and noticing the dialogue. Considering we know now that Savathun had her memories erased, probably had her Hive followers remember some about they're relationship-- they were their version of "The Last City". On first meeting, we killed the Hive guardian (probably no worries) but then [b]we killed the Hive ghost[/b]. And Savathun was referring to us as "honored guests" and not as usual 'guardians' but at the same time I think [b]she had sensed some Darkness in us[/b] because of Stasis and so we were strangers to her that came and attacked her people and killed a ghost-- something that "mirrors us" she said. These [b]Lucent Hive were pure Light[/b] already and we were half Light half Dark.
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We have to keep in mind [u][b]Savathun was basically "reset" into a fresh new guardian[/b][/u] and she wasn't who she was before and during season of the lost (much like Crow). Despite how devious Savathun's personality seems, she's still like a good leader supposedly trying to protect her people and the Traveler. I think in their eyes (and even objectively) we were the 'bad guys' in this one.
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In the lorebooks she would get the hive guardians rezzed and then immediately start torturing them until they were brainwashed into joining her cause. She was manipulative and evil, and even when she didn't have her memories she still continued to act like the witch queen, she'd come across as sympathetic to gain our trust, remember season of the lost?