"Every fairytale needs a good old fashioned villain." -Moriarty.
So, it sounds like their will be a lot of bad guys in Destiny. However we need something to make us hate the invading aliens, sure they destroyed most of humanity, but I get the impression that happened a little bit before the events of Destiny 1 take place.
What I'm saying is that we shouldn't be fighting a Hatfield and McCoy second generation war, our characters need a direct reason to stomp their faces in, and feel good afterwards.
Perhaps it will be something you discover, as you mop up a squad of Fallen, you're standing in a decades-old nursery, or maybe you'll discover bones of women and children in a bomb shelter. But if the game is going to have RPG elements it has to give proper incentive, otherwise the killing is just to enjoy the loot.
TLDR: Make us hate the aliens for more than just pushing us back, loot and the challenge of fighting is a good incentive, but we need to get a warm feeling by killing the enemy.
Any ideas on how Bungie should fuel the furnace of our Guardian's retribution?
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I like it. I know the Skyrim comparisons abound, but in Skyrim, almost every dungeon had a story, told through journals or the lay out of skeletons and dead bodies. I'm hoping every space in Destiny does the same thing, but like you said, at the same time imbuing this sense of devastation.