Honestly, I have no freaking idea. Girls are stupid. I put myself in the situation of saying "No, it's fine." When really, it's not. But at least if the male doesn't notice that it's not fine I don't take it out directly onto him because I know I'm stupid for saying it was fine in the first place. The trick with females is attention and petting in moderation. Very much like cats. Touch her where she doesn't want to be touched and your head will be bitten off.