The best feeling in all of Destiny is when you've thrown your knife, and it finds itself buried in the head of your target, killing it instantly. Instant smile, every freaking time.
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For me it was being level 3 and running into a warsat event having a bunch of other newbies show up and almost all die protecting it then all dancing and waving to each other as we headed our separate ways. Was the first day and a great feeling, it really set the tone for the beta. Fun.