They were born of humans in deep space under mysterious circumstances during the collapse.
Ghosts did not exists until the late Dark Age following the Collapse. The First Guardians didn't arrive until the very tail end and signaled the beginning of the City Age when they fought in wars to establish the Last City.
I've already explained this further below. Awoken existed and died long before the Ghosts began raising Guardians.
Lore remains intact.
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