That is exactly the kind of thing I figured was a major problem with implementing a lot of the features in this game but I really hope that bungie doesn't stick with that mindset going forward simply because it is EXTREMELY limiting.
I think the best way to go about it is add certain features that are next gen only. Don't make every feature next gen only but for instance, in some random free DLC or something, add in reverse levels, nightfall story missions, and lv 28 patrol for both old and new gen but also add 12 v 12 patrol pvp for new gen.
Everyone's getting new content while also being subtly pushed toward the newer consoles. They aren't required to enjoy the game, but to enjoy that mode.
Hell of a delicate balance though.
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Defiance is on 360: LOT of players there, perhaps it's just a server strength limitation. Speculation, speculation, speculation :-)